The 25 images shortlisted for Wildlife Photographer of the Year competition

Lumix People's Choice shortlist 2019 lets fans vote for their winner

Winner: Station squabble by Sam Rowley, UK. Rowley discovered the best way to photograph the mice inhabiting London’s Underground was to lie on the platform and wait. He only saw them fight over scraps of food dropped by passengers a few times, possibly because it is so abundant. This fight lasted a split second, before one grabbed a crumb and they went their separate ways. © Sam Rowley - Wildlife Photographer of the Year

Highly commended: Matching outfits by Michel Zoghzoghi, Lebanon. Zoghzoghi was in the Pantanal, Brazil photographing jaguars. One afternoon, as he was on the Tres Irmaos River, a mother and her cub crossed right in front of his boat. He watched mesmerised as they left the water holding an anaconda with a very similar pattern to their own. Michel Zoghzoghi / Wildlife Photographer of the Year

Highly commended: The surrogate mother by Martin Buzora, Canada. Elias Mugambi is a ranger at Lewa Wildlife Conservancy in northern Kenya. He often spends weeks away from his family caring for orphaned black rhinos like Kitui here. The young rhinos are in the sanctuary as a result of poaching or because their mothers are blind and cannot care for them safely in the wild. Martin Buzora / Wildlife Photographer of the Year

Highly commended: Losing the fight by Aaron Gekoski, UK. Orangutans have been used in degrading performances at Safari World, Bangkok – and many other locations – for decades. The shows were temporarily stopped in 2004 due to international pressure, but today the shows continue – twice a day, every day – with hundreds of people paying to watch the orangutans box, dance, play the drums and more. Aaron Gekoski / Wildlife Photographer of the Year

Highly commended: Spot the reindeer by Francis De Andres, Spain. The conditions for photographing at the Norwegian archipelago Svalbard are extreme, but wildlife has adapted to the environment and its freezing temperatures. De Andres found this composition of white arctic reindeer, which were observing him, both curious and charming. Francis De Andres / Wildlife Photographer of the Year

The unwelcome visitor by Salvador Colvee Nebot, Spain. Over several months, Nebot watched different species of bird use the dead flower spike of the agave in Valencia, Spain as a perch before descending to a small pond to drink. A pair of common kestrels were frequent visitors though each time they came magpies would hassle them. Salvador Colvee Nebot / Wildlife Photographer of the Year

Dressed for dawn, Csaba Tokolyi, Hungary. Tokolyi had been in a hide all night photographing nocturnal species and their activities, but as the golden light of dawn reflected on the surface of the water, an egret in wonderful breeding plumage stopped close by. The elongated scapular feathers covered the bird as if it was wearing a gown. Csaba Tökölyi / Wildlife Photographer of the Year

A pulsing sea by David Doubilet, USA. A school of red tooth triggerfish form a cloud of silhouettes above a river of convict blennies flowing over the coral in Verde Island Passage, Philippines. The Passage, a strait that separates the islands of Luzon and Mindoro, is one of the most productive marine ecosystems in the world. David Doubilet / Wildlife Photographer of the Year

Meeting place by Yaz Loukhal, France, After a rough journey by sea to the remote Snow Hill Island off the east coast of the Antarctic Penisula, Loukhal flew by helicopter and then trekked through thick snow to reach the emperor penguin colony. His efforts were rewarded with this incredible view of the whole colony. Yaz Loukhal / Wildlife Photographer of the Year

The humpback calf by Wayne Osborn, Australia. Osborn spotted this male humpback calf and its mother while diving off the Vava’u Island group in the Kingdom of Tonga. The calf kept a curious eye on Wayne as it twisted and turned before returning to its mother periodically to suckle. She was relaxed and motionless 20 metres (65 feet) below. Wayne Osborn / Wildlife Photographer of the Year

Big ears by Valeriy Maleev, Russia. Maleev was on a summer expedition to the Mongolian part of the Gobi Desert when he happened upon a long-eared jerboa. As blood moves through the ears of these usually nocturnal animals, excess heat dissipates across the skin and so the jerboa is able to stay cool. Valeriy Maleev / Wildlife Photographer of the Year

Trustful by Ingo Ardnt, Germany. For over two years Ardnt has followed the pumas of Torres del Paine National Park, in Patagonia, Chile. This female was so used to his presence that one day she fell asleep nearby. On awakening, she glanced at him in a familiar way, and he was able to capture this portrait of a completely relaxed puma. Ingo Ardnt / Wildlife Photographer of the Year

Training session by Stefan Christmann, Germany. When Christmann came across this penguin couple in Atka Bay, Antarctica, seemingly with an egg, he was surprised as it was too early in the season for egg laying. Upon closer inspection he discovered the egg was a snowball. Perhaps the diligent couple were practicing egg transfer in preparation for when their real egg arrived. This is possibly the first time it has ever been witnessed and documented. Stefan Christmann / Wildlife Photographer of the Year

What a poser by Clement Mwangi, Kenya. In Kenya’s Maasai Mara National Reserve, Mwangi spent time observing this beautiful leopard as she soaked up the last warm rays of the setting sun. Mwangi is mindful to remember to take pleasure in life’s simple moments – being all too aware that sometimes, as a wildlife photographer, you can miss the exceptional while looking for the unusual. Clement Mwangi / Wildlife Photographer of the Year

Mother knows best by Marion Volborn, Germany. While on a bear watching trip to the Nakina River in British Columbia, Canada Marion spotted a grizzly bear and her young cub approach a tree. The mother bear started to rub against the tree trunk and was followed shortly by the cub, imitating its mother. Marion Volborn / Wildlife Photographer of the Year

Tender play by Steve Levi, USA. It was early March and Levi spotted this mother polar bear and her two cubs after 10 days of looking. They had recently left their birthing den in Wapusk National Park, Canada, to begin the long journey to the sea ice so their mother could feed. After a nap the cubs were in a playful mood. Steve Levi / Wildlife Photographer of the Year

Teamwork by Jake Davis, USA. Jake was on a boat off the coast of Great Bear Rainforest, British Columbia, Canada where he watched humpback whales bubble-et feeding. Here the leader whale dives to locate the fish, once the fish are located, the rest of the pod swim in decreasing circles while blowing bubbles which create a net, trapping the fish. © Jake Davis/ Wildlife Photographer of the Year

A suitable gift by Marco Valentini, Italy. Valentini was in Hortobagyi National Park, Hungary when he spotted these kestrels displaying typical courtship behaviour. Here the female has just recovered an offering of a young green lizard from her suitor and in this touching moment she tenderly took hold of his claw. Marco Valentini / Wildlife Photographer of the Year

Beak to beak by Caludio Contreras Koob Mexico. Ria Lagartos Biosphere Reserve in the state of Yucatan is home to Mexico’s largest flock of Caribbean flamingos. This chick is less than five days old – it will stay in its nest less than a week before it joins a creche of other youngsters who wander around the colony searching for food. Claudio Contreras Koob / Wildlife Photographer of the Year

Inquisitive by Audan Rikardsen, Norway. From a hide on the coast of northern Norway, it took Rikardsen three years of planning to capture this majestic bird of prey in its coastal environment. After some time, the golden eagle became curious of the camera and seemed to like being in the spotlight. Audun Rikardsen / Wildlife Photographer of the Year

Ocean’s signature by Angel Fitor, Spain. Angel took this image in the waters off of Alicante, Spain. Immersed in a strong current, an otherwise slightly undulating salp chain twists and turning forming whimsical shapes. Salps move by contracting, which pumps water through their gelatinous bodies. Angel Fitor / Wildlife Photographer of the Year

Family get-together by Michael Schober, Austria. Marmots have become accustomed to the presence of humans in Hohe Tauern National Park, Austria and allow people to observe and photograph them at close range. This behaviour is beneficial for the marmots, as human company deters predators such as golden eagles. Michael Schober / Wildlife Photographer of the Year

Bon appetit by Lucas Bustamente, Ecuador. Night hikes through the Ecuadorian jungle are one of Bustamente’s favourite activities. With a keen interest in herpetology, he was overjoyed to spot this labiated rain frog which are abundant in the region. It had just caught a baby tarantula and its comical expression said ‘caught in the act!’ Lucas Bustamente / Wildlife Photographer of the Year

Winter’s tale by Valeriy Maleev, Russia. Valeriy encountered this Pallas’s cat while it was out hunting in the Mongolian grasslands – it was -42°C (-44°F) on that frosty day, but the fairy tale scene cancelled out the cold. Pallas’s cats are no bigger than a domestic cat and they stalk small rodents, birds and occasionally insects. © Valeriy Maleev - Wildlife Photographer of the Year 2

Captive by Marcus Westberg, Sweden. A giant panda sits in its cage in a breeding centre in Shaanxi, China. With a growing wild population and no realistic plan of how to breed and raise pandas for rerelease into the wild rather than a life in captivity – not to mention lack of habitat being the largest barrier to the continued spread of the wild population – it is unclear how such centres will benefit the species. © Marcus Westberg - Wildlife Photographer of the Year

The shortlist for Lumix's People's Choice Wildlife Photographer of the Year has been announced, and you can vote for the winner.

From more than 48,000 submissions from 100 countries, London's Natural History Museum has trimmed the list down to just 25. The photos in the running include Steve Levi's image titled Tender, portraying a family of polar bears; Aaron Gekosk's Losing the Fight, featuring a dressed up orangutan; Sam Rowley's image of fighting mice on the London Underground and Valeriy Maleev's photograph of a long-eared jerboa.

Click through the gallery above to see the full shortlist of photographs. 

In the shortlist there are photographers from 16 countries, including Lebanon, Mexico and Australia.

Voting is open until 2pm (GMT), Tuesday, February 4, and an exhibition of the shortlisted images is currently on display at the Natural History Museum. The display will be open until Sunday, May 31.

Wildlife Photographer of the Year is the longest-running photography competition in the world. Tim Littlewood, executive director of science at the Natural History Museum, said of the prize: "The Lumix People’s Choice images capture the essence of the competition; they all ignite a reaction about the natural world and make you see it differently.

"Showcasing breath-taking beauty, compassion and cruelty, it is impossible not to be moved by them," he added.