The World Nature Photography Awards winners - in pictures
World Nature Photographer of the Year grand prize winner and winner of the gold award in the Behaviour - Birds category: Two gannets under the water in the Shetland Islands, taken by Tracey Lund. All photos: World Nature Photography Awards
Gold award winner in the Behaviour - Amphibians and Reptiles category: A lava lizard standing on a marine iguana on the Galapagos Islands, taken by John Seagar
Gold award winner in the Underwater category: A female orca splitting a herring bait ball in Norway, taken by Andy Schmid
Gold award winner in the Animals in their Habitat category: Nubian Ibex in the Negev desert, Israel, taken by Amit Eshel
Gold award winner in the Planet Earth's Landscapes and Environments category: Beach and Vestrahorn mountain in Iceland, taken by Ivan Pedretti
Gold award winner in the Animal Portraits category: A black frogfish in Australia, taken by Nicolas Remy
Gold award winner in the Behaviour - Mammals category: Mother zebra and her foal being attacked by a cheetah in Kenya, taken by Alexander Brackx
Gold award winner in the Nature Photojournalism category: A sea lion pup in Baja California, Mexico, taken by Celia Kujala
Gold award winner in the Black and White Photo category: A leopard in East Rift Valley, Kenya, taken by Richard Li
Winner of the gold award in the Plants and Fungi category: Thismia thaithongiana in Thailand taken by Chatree Lertsintanakorn
Gold award winner in the People and Nature category: Traditional stilt fishermen try their luck at sunset in Sri Lanka, taken by Pavlos Evangelidis
Gold award winner in the Urban Wildlife category: A female sunbird in Israel, taken by Roy Wiesner
Gold award winner in the Nature Art category: Blue watery veins, glacial braids and golden sediment in Iceland, taken by Miki Spitzer
Gold award winner in the Behaviour - Invertebrates category: Sally Lightfoot crabs in the Galapagos Islands, taken by Bill Klipp